CAIRO: Saudi-led airstrikes killed at least 80 people near Yemen’s border with Saudi Arabia and in the capital Sanaa Wednesday, residents said, the deadliest day of bombing in over two months of war in Yemen.

Iran-backed Houthi rebels seized Sanaa last September and then thrust into central and south Yemen. Seeing the Houthi advance as a bridgehead for Iranian influence in the region, a Saudi-led coalition began airstrikes on March 26 in a campaign to restore Yemeni President Abed Rabbou Mansour Hadi to power.

Wednesday’s air raids on the Bakeel al-Meer area in Al-Hajjah province across Saudi Arabia’s border with Yemen killed at least 40 people, most of them civilians, local inhabitants said.

Tribesmen aligned with the Houthis have been fighting Saudi ground forces in the area, and border clashes have escalated the conflict between the Shiite Muslim rebels and the coalition of Sunni Muslim Gulf Arab states.

“Houthi gunmen were attacking Saudi border positions from this area but the coalition’s planes failed to hit the fighters and bombed civilians [instead],” a resident told Reuters by phone.Several hours later, airstrikes hit a special forces base allied with the Houthis in central Sanaa, the Houthi-run state news agency Saba said, in an account confirmed by residents.

“Around 40 people were martyred and more than 100 others were wounded, according to a preliminary toll, in bombing by the Saudi aggression’s planes on the Sabaaeen area in the capital Sanaa today,” the Saba dispatch said.

A Yemeni soldier who survived the attack said the raid hit a warehouse where soldiers and Houthi militiamen were receiving their weapons. “So far we’re not sure how many are dead,” the soldier told Reuters. “There were many people at the entrance to the warehouse, getting their weapons, farmers, cooks [and also soldiers]; these poor people were standing at the entrance to the warehouse. Two strikes, two hits, [occurred] one right after the other. The warehouses are completely destroyed.”

Yemeni army units loyal to former president Ali Abdullah Saleh, forced from power by a popular uprising in 2011, are fighting alongside the Houthis.

Arab warplanes and ships also hammered Yemen’s largest military port in the Red Sea city of Hudaida at dawn Wednesday, a local official said, the most serious attack on the country’s navy in over two months of war.

Hudaida and its military bases are aligned with the Houthis, the most powerful force in Yemen’s complex conflict, which also involves southern secessionist militias, local tribal forces and Islamist militants such as Al-Qaeda’s regional wing.

“The naval base was bombed by aircraft and ships. Large parts of it were destroyed and two warships were hit, and one of them, named the Bilqis, was destroyed and sank onto its side, and five gunboats shelled the administrative buildings of the base,” the official told Reuters by phone from Hudaida.

Houthi forces shelled the southern city of Aden, a bastion of resistance against their moves into Yemen’s south, and local fighters built on gains against the Houthis in recent days by seizing their last military post in the nearby city of Daleh.

Sunni Arab states see the Houthis as a proxy for projecting the power of archrival Iran in the Arabian Peninsula. Yemen’s exiled government in Saudi Arabia has said the Houthis must recognize its authority and vacate Yemen’s main cities before any peace talks can begin.

The United Nations said Tuesday that U.N.-backed negotiations which were set for May 28 in Geneva had been postponed.

In a new report Wednesday, World Health Organization chief Margaret Chan said that Yemen’s conflict has killed up to 2,000 people and wounded 8,000, including hundreds of women and children. She did not specify how many of the dead were civilians.

Recent U.N. estimates have said that at least 1,037 civilians, including 130 women and 234 children, have been killed in the fighting.

Chan also said that the killings sometimes included whole families, giving the example of a 65-year-old woman named Fathiya who lost 13 members of her family in an attack that left her the only guardian of three surviving grandchildren.

Journalists have also been a target following the Houthis seizing of Sanaa in September, taking over government institutions and ministries.

In the most recent incident, two young Yemeni journalists, Abdullah Qabil and Youssef al-Ayzari, were found dead after Houthis detained them while covering fighting in the city of Dhamar, south of Sanaa, according to the country’s Press Syndicate. The syndicate said the two were led by Houthis Wednesday to a site struck by Saudi-led airstrikes.